Luke Custer

Luke Custer's powers and abilities

Luke has various forms and abilities of beasts due to the chips he carries around.

Werewolf

Luke's first anthropomorphic form. It gives him razor-sharp claws to tear through preys (which can be charged up after arranging proper genetic modification), superhuman speed, howl that can stun low-level monsters, and enhanced jumping power.

Berserker Rage - a super-charged rush attack that annihilates all enemies standing in front of the werewolf

Rising Claw - an upper claw attack that sends enemies flying

Merman

⬜1 → ⬜2 → ⬜3 → ⬜4 → ⬜5

Luke's second anthropomorphic form. He can swim and breathe underwater. This form enables Luke to shoot air bubble bullets which are made by compressing airs within his own body, from his mouth. Merman can launch his own image made up of energy as a decoy to distract monsters, and can shoot Depthcharge, a chargeable energy bomb capable of breaking through certain impassable gates. In underwater Luke will be forced to progress using this form.

Wendigo

Luke's third anthropomorphic form. This form is designed to operate in extremely cold regions. It gives Luke superhuman strength, able to breathe blizzard that can instantly freeze victims to death, can create ice cubes which can be used as projectiles, and immunity to cold temperature. However, being a big creature created for power at the expense of maneuverability and agility, it is sluggish and that is this form's biggest drawback.

Garuda

⬜1 → ⬜2 → ⬜3 → ⬜4 → ⬜5

Luke's fourth anthropomorphic form. This beast form is designed for long-range flight operations. This form grants Luke an ability to fly. With this, Luke can go anywhere and can reach most of (save few ones that are underwater) secret or hidden bonus zones that were previously out of his reach. He can shoot feathers as if they are darts at enemies with lethal effect, can create a powerful tornado that devastates and annihilates all enemies foolish enough to run into it, and can swoop down on enemies for high damage. In some stages which require flying above Genome Mist, Luke will be forced to venture using Garuda form. Its only weakness is that it cannot walk.

Minotaur

Luke's fifth anthropomorphic form. This form is a giant humanoid bull made of fire and metal designed primarily for defense. This form grants Luke immunity to extremely high temperature and fire. In addition to goring his enemies with horns and punching his enemies with fiery fists, it can breathe flames that can incinerate enemies from its nose and can temporarily harden its body into a super armor which can repel even the largest known shells and bombs. When Minotaur changes himself into this steel form in mid-air, it falls down fast like a rock, crushing any enemies under it. Minotaur can also unleash a huge fiery explosion from his own body as a short range all-around attack, damaging any enemies nearby. Minotaur form also grants Luke superhuman endurance and resilience; as it takes lesser damages than other beast forms when hit, and when Minotaur takes hits he does not get knocked down to the ground like other beasts, but instead bounces backward, ready to launch immediate counterattack. His bull rush attack is immune to minor monster's attacks and is capable of mowing down any lesser monsters unintelligent enough to be in the way of Minotaur. Minotaur's speed accelerates further the longer Minotaur performs its bull rush. So long as he doesn't run into solid object, Minotaur can perform bull rush as long as it wants to.

Dragon

⬜1 → ⬜2 → ⬜3 → ⬜4 → ⬜5

This beast form is a dragon. It is the most powerful beast and each of its attacks leaves huge swath of destruction and corpses of monsters. Within the body of dragon lies a powerful powerhouse that enables dragon to constantly create electricity, which dragon utilizes as offensive weapon, on its own. His abilities involve in hurling lightning bombs at opponents, unleashing wide lightning sweep from his electrically charged that can annihilate any enemies nearby, and shoot highly concentrated lightning forward attack at frontal enemies (can only be used when fully charged so this should be used with caution). Although possessing wings, dragon only hovers so it doesn't fly the way Garuda does. And despite being the most powerful beast out of all forms, it is slow in both the movement and in attacks, leaving dragon somewhat vulnerable to enemies at times.